雅思阅读中的那些鲸鱼
Sperm whale skeleton.Richard Lydekker,1894.
The whale ear has specific adaptations to the marine environment.In
humans,the middle ear works as an impedance equalizer between the
outside air's low impedance and the cochlear fluid's high impedance.In
whales,and other marine mammals,there is no great difference between the
outer and inner environments.Instead of sound passing through the outer
ear to the middle ear,whales receive sound through the throat,from
which it passes through a low-impedance fat-filled cavity to the inner
ear.[41]The whale ear is acoustically isolated from the skull by
air-filled sinus pockets,which allow for greater directional hearing
underwater.[42]Odontocetes send out high frequency clicks from an organ
known as a melon.This melon consists of fat,and the skull of any such
creature containing a melon will have a large depression.The melon size
varies between species,the bigger the more dependent they are of it.A
beaked whale for example has a small bulge sitting on top of its
skull,whereas a sperm whale's head is filled up mainly with the
melon.[43][44][45][46]
The whale eye is relatively small for
its size,yet they do retain a good degree of eyesight.As well as
this,the eyes of a whale are placed on the sides of its head,so their
vision consists of two fields,rather than a binocular view like humans
have.When belugas surface,their lens and cornea correct the
nearsightedness that results from the refraction of light;they contain
both rod and cone cells,meaning they can see in both dim and bright
light,but they have far more rod cells than they do cone cells.Whales
do,however,lack short wavelength sensitive visual pigments in their cone
cells indicating a more limited capacity for colour vision than most
mammals.[47]Most whales have slightly flattened eyeballs,enlarged
pupils(which shrink as they surface to prevent damage),slightly
flattened corneas and a tapetum lucidum;these adaptations allow for
large amounts of light to pass through the eye and,therefore,a very
clear image of the surrounding area.In water,a whale can see around 10.7
metres(35 ft)ahead of itself,[citation needed][need quotation to
verify]but,of course,they have a smaller range above water.They also
have glands on the eyelids and outer corneal layer that act as
protection for the cornea.[48][49]
The olfactory lobes are
absent in toothed whales,suggesting that they have no sense of
smell.Some whales,such as the bowhead whale,possess a vomeronasal
organ,which does mean that they can"sniff out"krill.[50]
Whales are not thought to have a good sense of taste,as their taste
buds are atrophied or missing altogether.However,some toothed whales
have preferences between different kinds of fish,indicating some sort of
attachment to taste.The presence of the Jacobson's organ indicates that
whales can smell food once inside their mouth,which might be similar to
the sensation of taste.[51]
Communication
Whale vocalization is likely to serve several purposes.Some
species,such as the humpback whale,communicate using melodic
sounds,known as whale song.These sounds may be extremely loud,depending
on the species.Humpback whales only have been heard making clicks,while
toothed whales use sonar that may generate up to 20,000 watts of
sound(+73 dBm or+43 dBw)[52]and be heard for many miles.
Captive whales have occasionally been known to mimic human
speech.Scientists have suggested this indicates a strong desire on
behalf of the whales to communicate with humans,as whales have a very
different vocal mechanism,so imitating human speech likely takes
considerable effort.[53]
Whales emit two distinct kinds of
acoustic signals,which are called whistles and clicks:[54]Clicks are
quick broadband burst pulses,used for sonar,although some
lower-frequency broadband vocalizations may serve a non-echolocative
purpose such as communication;for example,the pulsed calls of
belugas.Pulses in a click train are emitted at intervals of~35–50
milliseconds,and in general these inter-click intervals are slightly
greater than the round-trip time of sound to the target.Whistles are
narrow-band frequency modulated(FM)signals,used for communicative
purposes,such as contact calls.
Intelligence
Whales are known to teach,learn,cooperate,scheme,and grieve.[55]The
neocortex of many species of whale is home to elongated spindle neurons
that,prior to 2007,were known only in hominids.[56]In humans,these cells
are involved in social conduct,emotions,judgement,and theory of
mind.Whale spindle neurons are found in areas of the brain that are
homologous to where they are found in humans,suggesting that they
perform a similar function.[57]
雅思阅读中的那些鲸鱼
Bubble net feeding
Brain size was previously considered a major indicator of the
intelligence of an animal.Since most of the brain is used for
maintaining bodily functions,greater ratios of brain to body mass may
increase the amount of brain mass available for more complex cognitive
tasks.Allometric analysis indicates that mammalian brain size scales at
approximately the?or?exponent of the body mass.Comparison of a
particular animal's brain size with the expected brain size based on
such allometric analysis provides an encephalisation quotient that can
be used as another indication of animal intelligence.Sperm whales have
the largest brain mass of any animal on earth,averaging 8,000 cubic
centimetres(490 in3)and 7.8 kilograms(17 lb)in mature males,in
comparison to the average human brain which averages 1,450 cubic
centimetres(88 in3)in mature males.[58]The brain to body mass ratio in
some odontocetes,such as belugas and narwhals,is second only to
humans.[59]
Small whales are known to engage in complex play
behaviour,which includes such things as producing stable underwater
toroidal air-core vortex rings or"bubble rings".There are two main
methods of bubble ring production:rapid puffing of a burst of air into
the water and allowing it to rise to the surface,forming a ring,or
swimming repeatedly in a circle and then stopping to inject air into the
helical vortex currents thus formed.They also appear to enjoy biting
the vortex-rings,so that they burst into many separate bubbles and then
rise quickly to the surface.[60]Some believe this is a means of
communication.[61]Whales are also known to produce bubble-nets for the
purpose of foraging.[62]
雅思阅读中的那些鲸鱼
A southern right whale sailing
Larger whales are also thought,to some degree,to engage in play.The
southern right whale,for example,elevates their tail fluke above the
water,remaining in the same position for a considerable amount of
time.This is known as"sailing".It appears to be a form of play and is
most commonly seen off the coast of Argentina and South Africa.Humpback
whales,among others,are also known to display this behaviour.[63]
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